Diprotodontia is an order of about 155 species of marsupialmammals including the kangaroos, wallabies, possums, koala, wombats, and many others. Extinct diprotodonts include the rhinoceros-sized Diprotodon, and Thylacoleo, the so-called "marsupial lion".
Characteristics
Living diprotodonts are almost all herbivores, as were most of those that are now extinct. A few insectivorous and omnivorous diprotodonts are known, and the Potoridae are almost unique among vertebrates in being largely fungivorous, but these seem to have arisen as relatively recent adaptations from the mainstream herbivorous lifestyle. The extinct thylacoleonids are the only known group to have exhibited carnivory on a large scale. Diprotodonts are restricted to Australasia. The earliest known fossils date to the late Oligocene, but their genesis certainly lies earlier than this, as large gaps occur in Australia's fossil record, with virtually no fossil record at all in geologically active New Guinea. The great diversity of known Oligocene diprotodonts suggests the order began to diverge well beforehand. Many of the largest and least athletic diprotodonts became extinct when humans first arrived in Australia about 50,000 years ago. Their extinction possibly occurred as a direct result of hunting, but was more probably a result of widespread habitat changes brought about by human activities—notably the use of fire. Two key anatomical features, in combination, identify Diprotodontia. Members of the order are, first, "diprotodont" : they have a pair of large, procumbent incisors on the lower jaw, a common feature of many early groups of mammals and mammaliforms. The diprotodont jaw is short, usually with three pairs of upper incisors, and no lower canines. The second trait distinguishing diprotodonts is "syndactyly", a fusing of the second and third digits of the foot up to the base of the claws, which leaves the claws themselves separate. Digit five is usually absent, and digit four is often greatly enlarged. Syndactyly is not particularly common and is generally posited as an adaptation to assist in climbing. Many modern diprotodonts, however, are strictlyterrestrial, and have evolved further adaptations to their feet to better suit this lifestyle. This makes the history of the tree-kangaroos particularly convoluted: it appears that the animals were arboreal at some time in the far distant past, moving afterward to the ground—gaining long kangaroo-like feet in the process—before returning to the trees, where they further developed a shortening and broadening of the hind feet and a novel climbing method.
Fossil record
The earliest known fossil of Diprotodontia dates back to the Late Oligocene, and the earliest identifiable species is Hypsiprymnodon bartholomaii from the Early Miocene. In 2020 fossils of Mukupirna, meaning big bones was found. This is thought to be the ancestor of the Vombatiformes.
Classification
Until recently, only two suborders in Diprotodontia were noted: Vombatiformes which encompassed the wombats and koala and Phalangerida which contained all other families. Kirschet al. split the families into three suborders. In addition, the six Phalangeriformes families are split into two superfamilies. Order DIPROTODONTIA
