Macronutrients provide structural material and energy. Some of the structural material can be used to generate energy internally, though the net energy depends on such factors as absorption and digestive effort, which vary substantially from instance to instance. Vitamins, minerals, fiber, and water do not provide energy, but are required for other reasons. A third class dietary material, fiber, seems also to be required, for both mechanical and biochemical reasons, though the exact reasons remain unclear. Molecules of carbohydrates and fats consist of carbon, hydrogen, and oxygen atoms. Carbohydrates range from simple monosaccharides to complex polysaccharides. Fats are triglycerides, made of assorted fatty acidmonomers bound to glycerol backbone. Some fatty acids, but not all, are essential in the diet: they cannot be synthesized in the body. Protein molecules contain nitrogen atoms in addition to carbon, oxygen, and hydrogen. The fundamental components of protein are nitrogen-containing amino acids. Essential amino acids cannot be made by the animal. Some of the amino acids are convertible to glucose and can be used for energy production just as ordinary glucose. By breaking down existing protein, some glucose can be produced internally; the remaining amino acids are discarded, primarily as urea in urine. This occurs normally only during prolonged starvation. Other dietary substances found in plant foods are not identified as essential nutrients but appear to impact health in both positive and negative ways. Most foods contain a mix of some or all of the nutrient classes, together with other substances. Some nutrients can be stored internally, while others are required more or less continuously. Poor health can be caused by a lack of required nutrients or, in extreme cases, too much of a required nutrient. For example, both salt provides sodium and chloride, both essential nutrients, but will cause illness or even death in too large amounts. Dietary fibre is a carbohydrate that is incompletely absorbed in some animals.
Protein
Proteins are the basis of many animal body structures. They also form the enzymes which control chemical reactions throughout the body. Each molecule is composed of amino acids which are characterized by the inclusion of nitrogen and sometimes sulfur. The body requires amino acids to produce new proteins and to replace damaged proteins. As there is no protein or amino acid storage provision, amino acids must be present in the diet. Excess amino acids are discarded, typically in the urine. For all animals, some amino acids are essential and some are non-essential. A diet that contains adequate amounts of amino acids is particularly important in some situations: during early development and maturation, pregnancy, lactation, or injury. A few amino acids from protein can be converted into glucose and used for fuel through a process called gluconeogenesis; this is done in quantity only during starvation.
Minerals
Dietary minerals are the chemical elements required by living organisms, other than the four elements carbon, hydrogen, nitrogen, and oxygen that are present in nearly all organic molecules. The term "mineral" is archaic, since the intent is to describe simply the less common elements in the diet. Many elements are essential in relative quantity; they are usually called "bulk minerals". Some are structural, but many play a role as electrolytes. These include:
Sulfur for three amino acids and therefore many proteins
Many elements are required in trace amounts, usually because they play a catalytic role in enzymes.
Vitamins
deficiencies may result in disease conditions. Excess of some vitamins is also dangerous to health, and animal nutrition researchers have managed to establish safe levels for some common companion animals. Deficiency or excess of minerals can also have serious health consequences. ASH Though not a nutrient as such, an entry for ash is sometimes found on nutrition labels, especially for pet food. This entry measures the weight of inorganic material left over after the food is burned for two hours at 600 °C. Thus, it does not include water, fibre, and nutrients that provide calories, but it does include some nutrients, such as minerals Too much ash may contribute to feline urological syndrome in domestic cats.
